Botany

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Botany is the scientific study of plants. As a branch of biology, it is also called plant science(s), phytology, or plant biology. Botany covers a wide range of scientific disciplines that study plants, algae, and fungi including: structure, growth, reproduction, metabolism, development, diseases, and chemical properties and evolutionary relationships between the different groups.

The study of plants and botany began with tribal lore, used to identify edible, medicinal and poisonous plants, making botany one of the oldest sciences. From this ancient interest in plants, the scope of botany has increased to include the study of over 550,000 kinds or species of living organisms.

Scope and importance of botany

Hibiscus

As with other life forms in biology, plant life can be studied from different perspectives, from the molecular, genetic and biochemical level through organelles, cells, tissues, organs, individuals, plant populations, and communities of plants. At each of these levels a botanist might be concerned with the classification (taxonomy), structure (anatomy), or function (physiology) of plant life.

Historically, botany covers all organisms that were not considered to be animals. Some of these "plant-like" organisms include fungi (studied in mycology), bacteria and viruses (studied in microbiology), and algae (studied in phycology). Most algae, fungi, and microbes are no longer considered to be in the plant kingdom. However, attention is still given to them by botanists, and bacteria, fungi, and algae are usually covered in introductory botany courses.

The study of plants has importance for a number of reasons. Plants are a fundamental part of life on earth. They generate the oxygen, food, fibres, fuel and medicine that allow higher life forms to exist. Plants also absorb carbon dioxide through photosynthesis, a minor greenhouse gas that in large amounts can effect global climate. It is believed that the evolution of plants has changed the global atmosphere of the earth early in the earth's history and paleobotanists study ancient plants in the fossil record. A good understanding of plants is crucial to the future of human societies as it allows us to:

Produce food to feed an expanding population
Understand fundamental life processes
Produce medicine and materials to treat diseases and other ailments
Understand environmental changes more clearly

Human nutrition

Nearly all the food we eat comes (directly and indirectly) from plants like this American long grain rice.

Virtually all foods eaten comes from plants, either directly from staple foods and other fruit and vegetables, or indirectly through livestock or other animals, which rely on plants for their nutrition. Plants are the fundamental base of nearly all food chains because they use the energy from the sun and nutrients from the soil and atmosphere and convert them into a form that can be consumed and utilized by animals, this is what ecologists call the first trophic level. Botanists also study how plants produce food we can eat and how to increase yields and therefore their work is important in mankind's ability to feed the world and provide food security for future generations, for example through plant breeding. Botanists also study weeds, plants which are considered to be a nuisance in a particular location. Weeds are a considerable problem in agriculture, and botany provides some of the basic science used to understand how to minimize 'weed' impact in agriculture and native ecosystems. Ethnobotany is the study of the relationships between plants and people.

Gregor Mendel laid the foundations of genetics from his studies of plants.

Fundamental life processes

Plants are convenient organisms in which fundamental life processes (like cell division and protein synthesis for example) can be studied, without the ethical dilemmas of studying animals or humans. The genetic laws of inheritance were discovered in this way by Gregor Mendel, who was studying the way pea shape is inherited. What Mendel learned from studying plants has had far reaching benefits outside of botany. Additionally, Barbara McClintock discovered 'jumping genes' by studying maize. These are a few examples that demonstrate how botanical research has an ongoing relevance to the understanding of fundamental biological processes.

Medicine and materials

Many medicinal and recreational drugs, like cannabis, caffeine, and nicotine come directly from the plant kingdom. Aspirin, which originally came from the bark of willow trees, is just one example. There may be many novel cures for diseases provided by plants, waiting to be discovered. Popular stimulants like coffee, chocolate, tobacco, and tea also come from plants. Most alcoholic beverages come from fermenting plants such as barley malt and grapes.

Plants also provide us with many natural materials, such as cotton, wood, paper, linen, vegetable oils, some types of rope, and rubber. The production of silk would not be possible without the cultivation of the mulberry plant. Sugarcane, rapeseed, soy and other plants with a highly-fermentable sugar or oil content have recently been put to use as sources of biofuels, which are important alternatives to fossil fuels, see biodiesel.

Environmental changes

Plants can also help us understand changes in on our environment in many ways.

Understanding habitat destruction and species extinction is dependent on an accurate and complete catalog of plant systematics and taxonomy.
Plant responses to ultraviolet radiation can help us monitor problems like the ozone depletion.
Analyzing pollen deposited by plants thousands or millions of years ago can help scientists to reconstruct past climates and predict future ones, an essential part of climate change research.
Recording and analyzing the timing of plant life cycles are important parts of phenology used in climate-change research.
Lichens, which are sensitive to atmospheric conditions, have been extensively used as pollution indicators.

In many different ways, plants can act a little like the 'miners canary', an early warning system alerting us to important changes in our environment. In addition to these practical and scientific reasons, plants are extremely valuable as recreation for millions of people who enjoy gardening, horticultural and culinary uses of plants every day.

Etymology

From Greek βοτάνη = "pasture, grass, fodder", perhaps via the idea of a livestock keeper needing to know which plants are safe for livestock to eat.

History

The traditional tools of a botanist.

Among the earliest of botanical works, written around 300 B.C., are two large treatises by Theophrastus: On the History of Plants (Historia Plantarum) and On the Causes of Plants. Together these books constitute the most important contribution to botanical science during antiquity and on into the Middle Ages. The Roman medical writer Dioscorides provides important evidence on Greek and Roman knowledge of medicinal plants.

In ancient China, the recorded listing of different plants and herb concoctions for pharmaceutical purposes spans back to at least the Warring States (481 BC-221 BC). Many Chinese writers over the centuries contributed to the written knowledge of herbal pharmaceutics. There was the Han Dynasty (202 BC-220 AD) written work of the Huangdi Neijing and the famous pharmacologist Zhang Ji of the 2nd century. There was also the 11th century scientists and statesmen Su Song and Shen Kuo, who compiled treatises on herbal medicine and included the use of mineralogy.

In 1665, using an early microscope, Robert Hooke discovered cells in cork, and a short time later in living plant tissue. The German Leonhart Fuchs, the Swiss Conrad von Gesner, and the British authors Nicholas Culpeper and John Gerard published herbals that gave information on the medicinal uses of plants.

In 1754 Carl von Linné (Carl Linnaeus) devided the plant Kingdom into 25 classes. One, the Cryptogamia, included all the plants with concealed reproductive parts (algae, fungi, mosses and liverworts and ferns).^[1]

Modern botany

A considerable amount of new knowledge today is being generated from studying model plants like Arabidopsis thaliana. This weedy species in the mustard family was one of the first plants to have its genome sequenced. The sequencing of the rice (Oryza sativa) genome and a large international research community have made rice the de facto cereal/grass/monocot model. Another grass species, Brachypodium distachyon is also emerging as an experimental model for understanding the genetic, cellular and molecular biology of temperate grasses. Other commercially-important staple foods like wheat, maize, barley, rye, pearl millet and soybean are also having their genomes sequenced. Some of these are challenging to sequence because they have more than two haploid (n) sets of chromosomes, a condition known as polyploidy, common in the plant kingdom. Chlamydomonas reinhardtii (a single-celled, green alga) is another plant model organism that has been extensively studied and provided important insights into cell biology.

In 1998 the Angiosperm Phylogeny Group published a phylogeny of flowering plants based on an analysis of DNA sequences from most families of flowering plants. As a result of this work, major questions such which families represent the earliest branches in the genealogy of angiosperms are now understood. Investigating how plant species are related to each other allows botanists to better understand the process of evolution in plants.

Subdisciplines of Botany

Agronomy—Application of plant science to crop production
Bryology—Mosses, liverworts, and hornwarts
Economic botany—The place of plants in economics
Ethnobotany—Relationship between humans and plants
Forestry—Forest management and related studies
Horticulture—Cultivated plants
Paleobotany—Fossil plants
Palynology—Pollen and spores
Phytochemistry—Plant secondary chemistry and chemical processes
Phytomorphology—Structure and life cycles
Plant anatomy—Cell and tissue structure
Plant ecology—Role of plants in the environment
Plant genetics—Genetic inheritance in plants
Plant pathology—Plant diseases
Plant systematics—Classification and naming of plants

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Botany

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